Hours after receiving an urgent call in his Parris apartment, the protagonist of D/Generation, a nameless courier, arrives in London via his Jet-Pack, touching down on the 80th floor of Genoq Biolabs' medical research facility with an important package for lead Genoq scientist Jean-Paul Derrida. As soon as he enters, however, the doors shut and seal themselves behind him. This, however, proves to be the least of his problems, as the receptionist soon informs him of Genoq's dire state. All lines of communication to the outside world have been severed, and furthermore the building's security systems and dangerous Neogens have run amok. Despite the perils within and the receptionist's adamant warnings, the courier chooses to proceed, perhaps believing that the item in his possession holds the key to stopping the chaos which grips Genoq.

Over the course of his journey into the building's upper reaches, the courier learns that Genoq had been steeped in the realm of illegal bioweapon development for some time, and that their medical research, which had long ago become unprofitable for them, was little more than a front for their more lucrative practice of creating and selling biological weapons called Neogens. Derrida had been brought on by the company to spearhead a project which would create the ultimate Neogen, and the fruit of his efforts, an artificial organism dubbed D/Generation, was ultimately the source of the disturbances within the building. Further complicating matters, he also learns that Genoq's corporate leaders have authorized an airstrike against the building in order to prevent details of their activities from going public.

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